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CIPK-B is essential for salt stress signalling in Marchantia polymorpha.

Tansley, C., et al. · 2023 · New Phytologist   research

doi:10.1111/nph.18633   PMID:36660914   PMC10953335

Linked genes (6)  1 core 5 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp2g26670 MpCIPK-B experimental subject CBL-interacting kinase in the AtCIPK24/SOS2 clade, the central subject of the study. Salt-downregulated, osmotic-upregulated; interacts with MpCBL-A/B in yeast and all MpCBLs in planta. CRISPR knockouts show increased salt sensitivity (chlorosis/death from 50 mM NaCl) but normal sorbitol tolerance, establishing it as essential for ionic salt-stress signalling.
Mp2g07750 MpCBL-A experimental tool Calcineurin B-like protein analyzed by phylogeny, expression, and interaction assays. Groups with tonoplast Arabidopsis CBLs but lacks the localization motif; downregulated by salt yet upregulated by osmotic stress; interacts with both MpCIPKs in planta.
Mp4g00900 MpCBL-B experimental tool Type I CBL with the MGCxxS/T plasma-membrane motif; modestly downregulated by salt and upregulated by osmotic stress; interacts with both MpCIPKs and Arabidopsis CIPK24 in yeast and in planta.
Mp5g19810 MpCBL-C experimental tool Type I CBL proposed as the SOS3/AtCBL4 homologue; strongly salt-downregulated but osmotic-stress-insensitive, suggesting an ionic-stress role; interacts with both MpCIPKs in planta.
Mp1g05680 MpCIPK-A experimental tool CBL-interacting kinase in a bryophyte-specific clade; salt-downregulated and osmotic-insensitive; binds all MpCBLs and Arabidopsis CBL4 in yeast and in planta, possibly via nonspecific interactions.
MpSOS1 missing sequence background Putative Na+/H+ antiporter; BLAST with AtSOS1 identifies two possible M. polymorpha homologues, proposed as downstream targets of MpCIPK-B, but phosphorylation/activation remains to be established.