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Marchantia polymorpha, a New Model Plant for Autophagy Studies.

Norizuki, T., et al. · 2019 · Frontiers in Plant Science   research

doi:10.3389/fpls.2019.00935   PMID:31379911   PMC6652269

Linked genes (7)  4 core 3 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp6g18570 MpATG2 experimental subject A core autophagy gene knocked out by CRISPR/Cas9 (Mpatg2-1ge frameshift). The mutant blocks vacuolar transport of fluorescent MpATG8, showing MpATG2 is required for autophagosome-mediated delivery and autophagy-dependent chlorophyll maintenance under nutrient stress.
Mp1g12840 MpATG5 experimental subject Core autophagy gene disrupted via CRISPR/Cas9 (Mpatg5-1ge). The mutant shows early senescence, blocked MpATG8 vacuolar transport, and starvation hypersensitivity, demonstrating its requirement for autophagosome formation in M. polymorpha.
Mp2g07850 MpATG7 experimental subject E1-like activating enzyme gene knocked out (Mpatg7-1ge frameshift via CRISPR/Cas9). The mutant fails to transport MpATG8 to the vacuole, shows reduced chlorophyll and early senescence, establishing MpATG7 as essential for autophagy under nutrient starvation.
MpATG8 missing experimental subject Ubiquitin-like autophagosome protein transported into the vacuole in an MpATG2-, MpATG5-, and MpATG7-dependent manner; exists as two isoforms (a/b). Fluorescently tagged MpATG8 serves as an autophagosome marker, accumulating in vacuolar autophagic bodies upon concanamycin A treatment.
Mp1g21590 MpATG8a experimental tool Isoform expressed as mCitrine-MpATG8a fusion under its native promoter; used as an autophagy marker showing autophagy-dependent vacuolar puncta and cleavage on immunoblots.
Mp5g05930 MpATG8b experimental tool Isoform expressed as mCitrine/mRFP-MpATG8b fusion; colocalizes with MpATG8a in the same autophagosomes/autophagic bodies, used as an interchangeable autophagosome marker.
MpATG18 missing sequence background Identified by comparative genomic and phylogenetic analysis; M. polymorpha harbors conventional and plant-unique (BCAS3-domain) ATG18 types, with the plant-unique form predating Streptophyta.