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H2A ubiquitination is essential for Polycomb Repressive Complex 1-mediated gene regulation in Marchantia polymorpha.

Liu, S., et al. · 2021b · Genome Biology   research

doi:10.1186/s13059-021-02476-y   PMID:34465381   PMC8408974

Linked genes (4)  3 core 1 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp7g12670 MpBMI1 experimental subject Core PRC1 RING-finger E3 ligase (Mp7g12670) identified by BLAST; CRISPR knockout combined with Mpbmi1l caused strongly reduced growth and gemma-cup defects, global H2Aub loss by western blot, and altered H2Aub/H3K27me3 by ChIP-seq, showing it deposits H2Aub for Polycomb gene repression.
Mp6g09730 MpBMI1L missing experimental subject Second BMI1 homolog (Mp6g09730) with only a RAWUL domain; functions redundantly with MpBMI1, as Mpbmi1/1l double knockouts were more severely affected than single mutants and lost H2Aub, establishing it as a PRC1 component required for H2A ubiquitination.
Mp3g02370 MpH2A experimental subject Single canonical H2A whose K115/K116/K119 were mutated to arginine via CRISPR; western blot confirmed reduced H2Aub, ChIP-seq/RNA-seq showed H2Aub required for H3K27me3 at genes and TEs with thallus and gemmae-dormancy phenotypes, demonstrating the three lysines are redundantly ubiquitinated in vivo.
MpRING missing citation comparator PRC1 catalytic component mentioned only in discussion, with possible BMI1-independent catalytic activity inferred from Arabidopsis AtRING1A/B.