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The wound- activated ERF15 transcription factor drives Marchantia polymorpha regeneration by activating an oxylipin biosynthesis feedback loop.

Liang, Y., et al. · 2022 · Science Advances   research

doi:10.1126/sciadv.abo7737   PMID:35960801   PMC9374346

Linked genes (15)  12 core 3 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp5g16260 MpAOS2 experimental subject Oxylipin biosynthesis gene (Mp5g16260) whose wound-induced transcript levels are MpERF15-dependent, being significantly lower in Mperf15ko.
Mp2g26760 MpASA1 missing experimental subject Auxin-biosynthesis (anthranilate synthase) gene acting downstream of MpERF15; its wound-induced expression is significantly reduced in Mperf15ko at 4 h post-wounding by RT-qPCR.
Mp2g26590 MpCOI1 experimental subject Oxylipin (COI1) receptor; MpERF15 acts both upstream and downstream of it. MpERF15 expression is strongly reduced in Mpcoi1-2 mutants, and MpERF15OE phenotypes are lost in the Mpcoi1-2 background, placing MpCOI1 in the feedback loop.
Mp7g06220 MpDIR17 experimental subject Allene oxide cyclase (Mp7g06220) catalyzing a key OPDA/dn-OPDA biosynthesis step. Mpaocko mutants regenerate slowly, a defect rescued by OPDA treatment; its wound-induced expression is reduced in Mperf15ko and Mpcoi1-2.
Mp5g16520 MpDIR38 experimental subject Dirigent-like protein gene (Mp5g16520), a previously known wound-inducible marker confirmed by RT-qPCR to be MpERF15-dependent.
Mp5g16470 MpDIR42 experimental subject Dirigent-like protein gene belonging to the wound-inducible, MpERF15-dependent gene cluster identified by RNA-seq.
MpDIR43 missing experimental subject Dirigent-like protein gene belonging to the wound-inducible, MpERF15-dependent gene cluster identified by RNA-seq.
Mp7g09350 MpERF15 experimental subject Wound-activated ERF transcription factor that is instantly induced after wounding and is essential for gemmaling regeneration. Knockouts (Mperf15ko) show regeneration defects while overexpression lines produce extra apical notches and ectopic meristems. Drives a positive oxylipin-biosynthesis feedback loop, operating both upstream and downstream of MpCOI1.
Mp5g06970 MpERF20 experimental subject MpERF20/LAXR (Mp5g06970), a late wound-inducible factor (peak ~6 h) acting downstream of MpERF15; wound inducibility partly depends on MpCOI1, MpAOC and MpERF15, and expression is elevated in MpERF15OE thalli.
Mp6g06230 MpJAZ experimental subject The single M. polymorpha JAZ repressor, phylogenetically closest to subgroup V; shown by yeast two-hybrid to interact with MpERF15 (and with Arabidopsis JAZ3/4/9).
Mp2g00660 MpLOX1 experimental subject Lipoxygenase, wound-inducible in an MpERF15-dependent manner and induced in MpERF15OE lines; likely acts in the dn-OPDA biosynthetic pathway.
Mp5g21680 MpLOX4 experimental subject Lipoxygenase, wound-inducible in an MpERF15-dependent manner; repressed in MpERF15OE lines and likely active in the OPDA biosynthetic pathway.
Mp6g11010 MpACTIN7 experimental tool Constitutive actin gene (Mp6g11010) used as a reference gene for RT-qPCR normalization.
Mp3g23400 MpEF1 sequence tool Constitutive ELONGATION FACTOR 1alpha promoter (Mp3g23400) used to drive overexpression constructs and as an RT-qPCR reference gene.
Mp8g08780 MpYUC2 experimental tool Used as a proMpYUC2:GUS reporter line to mark apical notches/meristematic activity, with more GUS dots in MpERF15OE than WT.