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R2R3-MYB transcription factor GEMMA CUP-ASSOCIATED MYB1 mediates the cytokinin signal to achieve proper organ development in Marchantia polymorpha.

Aki, S. S., et al. · 2022 · Scientific Reports   research

doi:10.1038/s41598-022-25684-3   PMID:36477255   PMC9729187

Linked genes (10)  3 core 7 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp6g04830 MpGCAM1 experimental subject R2R3-MYB transcription factor functioning downstream of cytokinin signaling; knockout mutants show defective gemma cup formation while inducible overexpression yields undifferentiated cell clumps. Indirectly upregulated by MpRRB-mediated signaling (not a direct target) and requires de novo protein synthesis for induction.
Mp3g03810 MpRRA experimental subject Type-A response regulator acting as a negative regulator of cytokinin signaling; knockout enhances cytokinin signaling and increases gemma cup formation. Cytokinin-induced in wild type but not in Mprrb, and the Mpgcam1 Mprra double mutant produces curled thalli without gemma cups.
Mp4g20600 MpRRB experimental subject Type-B response regulator central to cytokinin signaling; knockout lines show defective gemma cup formation and increased rhizoids. Indirectly upregulates MpGCAM1 and drives expression of nitrogen-transport and auxin-related genes; active at the apical notch of young thalli.
Mp5g10910 MpCKX2 experimental tool Cytokinin oxidase whose overexpression reduces endogenous cytokinins, used to lower cytokinin levels and assay MpGCAM1 expression (proMpGCAM1:GUS); overexpressing lines show fewer/no gemmiparous cells and reduced MpGCAM1 transcripts.
Mp3g23400 MpEF1 experimental tool Elongation factor 1-alpha used as the constitutive reference gene to normalize mRNA levels in qRT-PCR analyses.
Mp5g03090 MpCKX1 sequence background One of two cytokinin oxidase genes in M. polymorpha; listed among upregulated genes in RNA-seq but not functionally characterized here.
Mp2g14010 MpGH3B experimental background Acyl acid amido synthetase gene downregulated by MpRRB-mediated cytokinin signaling, indicating adjustment of auxin homeostasis.
Mp1g16480 MpIPT1 sequence background One of two cytokinin biosynthesis genes; not upregulated by MpGCAM1 overexpression, mentioned as background to cytokinin biosynthesis.
Mp8g02960 MpIPT2 sequence background Cytokinin biosynthesis gene orthologous to Arabidopsis IPT9; downregulated under MpRRB-mediated signaling and not upregulated by MpGCAM1 overexpression.
Mp6g07950 MpPIN3 experimental background Auxin efflux carrier gene induced by MpRRB-mediated cytokinin signaling, cited as evidence of cytokinin-auxin cross-talk.