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Biosynthesis of gibberellin-related compounds modulates far-red light responses in the liverwort Marchantia polymorpha.

Sun, R., et al. · 2023 · Plant Cell   research

doi:10.1093/plcell/koad216   PMID:37597168   PMC10615216

Linked genes (21)  9 core 12 peripheral

Gene ID Name Evidence / Role Function (this paper)
MpDTPS3 missing experimental subject Diterpene synthase 3 (= MpCPS); ent-copalyl diphosphate synthase in GA-related biosynthesis.
MpDTPS4 missing experimental subject Diterpene synthase 4 (= MpKS); kaurene synthase-type in GA-related biosynthesis.
Mp4g23680 MpKAOL1 experimental subject Loss-of-function mutant (Mp_kaol1ge) generated and assayed in yeast. Yeast expressing MpKAOL1 consumed KA and produced GA12, identifying it as the active KAO downstream of KA in GAMp biosynthesis.
Mp1g25410 MpKAOL2 experimental subject KAO-like cytochrome P450 (Mp1g25410); gibberellin-related diterpenoid biosynthesis modulating far-red responses.
Mp3g18320 MpKOL1 experimental subject Loss-of-function mutant Mp_kol1ld generated and enzyme assayed in yeast. Yeast expressing MpKOL1 consumed ent-kaurenoic acid (KA) and produced GA12; mutant showed enlarged thalli under cW+cFR, identifying MpKOL1 as the active KO required for FR response.
Mp3g17350 MpPIF experimental subject Knockout Mp_pifko analyzed by transcriptome; encodes a PIF transcription factor. FR-induced up-regulation of biosynthesis genes (e.g. MpKOL1) occurred only in wild-type and not Mp_pifko, showing MpPIF mediates FR responses.
Mp6g05430 MpTPS1 experimental subject Loss-of-function mutant Mp_tps1ld generated; minor ent-kaurene producer. Mutant morphology was similar to wild-type under cW and cW+cFR, indicating MpTPS1 is not a major gene required for GAMp biosynthesis.
Mp2g07200 MpTPS3 experimental subject CRISPR large-deletion mutants (Mp_cpsld) were generated; encodes the first enzyme of GA-related diterpenoid biosynthesis. Under cW+cFR, mutant thalli were significantly larger than wild-type, showing MpCPS is required for far-red-light developmental responses (hyponasty, slender thallus).
Mp6g05950 MpTPS4 experimental subject Loss-of-function mutant Mp_ksld was generated; acts downstream of MpCPS producing ent-kaurene. Mutant phenotype under cW+cFR resembled Mp_cpsld with significantly larger thalli, indicating MpKS is required for the FR response.
Mp2g10420 MpKAOL3 experimental comparator Homolog whose complete-deletion mutant Mp_kaol3ld produced no GA12 and showed no severe FR-response defect, lacking MpKAOL1's activity.
Mp2g01950 MpKOL2 experimental comparator Homolog tested in yeast; produced no KA and its mutant showed no FR-response phenotype, so it lacks the KA-producing activity of MpKOL1.
Mp2g01940 MpKOL3 experimental comparator Homolog tested in yeast; produced no KA and its mutant showed no FR-response phenotype, lacking MpKOL1's activity.
Mp3g23300 MpBNB experimental tool Citrine knock-in line (MpBNB-Cit) used as a marker; MpBONOBO protein accumulated in initial cells and immature gametangia to visualize early gametangium differentiation.
Mp5g19490 MpABA4 experimental background DEG: ABA metabolic gene induced by FR enrichment.
Mp5g10910 MpCKX2 experimental background DEG: cytokinin deactivation gene (CYTOKININ OXIDASE2) down-regulated in Mp_cps-4ld.
Mp2g25940 MpCYP707A experimental background DEG: ABA metabolic gene altered by FR enrichment.
Mp5g20660 MpGRAS6 citation background Cited as a GA-signaling repressor; its overexpression inhibits MpPIF-mediated FR responses, referenced to distinguish GAMp signaling.
Mp8g02960 MpIPT2 experimental background DEG: cytokinin biosynthesis gene (ISOPENTENYLTRANSFERASE2) down-regulated in Mp_cps-4ld.
Mp1g00270 MpLOG experimental background DEG: cytokinin-activating gene (LONELY GUY) down-regulated in Mp_cps-4ld.
Mp2g07800 MpNCED experimental background DEG: ABA biosynthesis gene induced by FR enrichment.
Mp3g13150 MpTPS6 experimental background DEG: expression encoding a cis-kolavenol-producing enzyme was significantly decreased in Mp_cps-4ld.