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The cAMP signaling module regulates sperm motility in the liverwort Marchantia polymorpha.

Yamamoto, C., et al. · 2024 · Proceeding of the National Academy of Sciences USA   research

doi:10.1073/pnas.2322211121   PMID:38593080   PMC11032487

Linked genes (6)  3 core 3 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp7g08500 MpCAPE experimental subject Dual adenylyl cyclase/phosphodiesterase that synthesizes and hydrolyzes cAMP, expressed during maturation of male reproductive organs. Knockout mutants are male-infertile with impaired flagellar motility, swimming in circles instead of straight (16.4 vs 43.5 um/s WT). Complementation restored normal motility, identifying it as the central cAMP regulator of sperm motility.
Mp2g09970 MpPKAR1 experimental subject PKA regulatory subunit acting as a downstream cAMP effector, highly expressed in antheridia. Double-knockout with MpPKAR2 caused abnormal/trembling sperm motility and reduced swimming linearity and speed comparable to CAPE mutants. Complementation of MpPKAR1 restored normal motility.
Mp8g17350 MpPKAR2 experimental subject PKA regulatory subunit and cAMP effector, highly expressed in antheridia. MpPKAR1/MpPKAR2 double-knockout mutants showed trembling/immotile sperm with defects in swimming linearity and speed, demonstrating PKA function downstream of cAMP in sperm motility.
Mp3g23400 MpEF1 citation tool Elongation factor used as an internal reference for normalizing transcript levels in expression heat maps.
MpPKAC sequence background PKA catalytic subunit identified by CNB domain search; conserved PKA-C inhibitor-site signature aligning with mammalian PKA-Calpha/beta, not experimentally tested.
Mp6g02750 MpTIR1 citation background Auxin receptor ortholog cited as an alternative cAMP source but ruled out, since it acts in vegetative tissues not male organs.