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Spatial distribution of isoprenoid enzymes and MpABCG1 transporter influences sesquiterpene accumulation in Marchantia polymorpha oil bodies.

Forestier ECF, Asprilla P, Bonter I, Romani F, Frangedakis E, Haseloff J. · 2026 · Communications biology   research

doi:10.1038/s42003-025-09508-4   PMID:41772152   PMC13068942

Linked genes (30)  13 core 17 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp8g13070 MpABCG1 experimental subject Central subject of the paper. MpABCG1 is an ABCG-family transporter localized at the oil body membrane; CRISPR knockout dramatically reduced sesquiterpene accumulation in oil bodies, while overexpression mildly increased endogenous sesquiterpene levels, indicating MpABCG1 mediates sesquiterpene transport/accumulation into oil bodies.
Mp6g04420 MpDXR1 experimental subject MEP pathway step-2 enzyme whose localization was characterized in oil body cells in this work.
Mp8g01240 MpDXR2 experimental subject DXR isoform reported as oil body-specific in expression/localization within this study.
Mp2g09130 MpDXS1 experimental subject MEP pathway entry enzyme studied for subcellular localization; MpDXS1 localized to plastids with enhanced expression in oil body cells.
Mp4g14720 MpDXS2 experimental subject Second DXS isoform; localized to plastids and associated with oil body cell isoprenoid biosynthesis in this study.
MpFPS missing experimental subject Farnesyl pyrophosphate synthase (corresponding to IDS1) producing the sesquiterpene precursor FPP; shown to be cytosolic in oil body cells, consistent with cytosolic sesquiterpene biosynthesis.
Mp6g04320 MpFTPSL2 experimental subject Endogenous sesquiterpene synthase (fungal terpene synthase-like 2) previously characterized; in this paper preliminary localization evidence places it partly in the oil body lumen, consistent with sesquiterpene synthesis associated with oil bodies.
MpGGPPS missing experimental subject Geranylgeranyl pyrophosphate synthase (re-annotated from MpIDS2) producing the diterpene precursor; characterized as plastid-localized in this study.
Mp5g09880 MpHDR experimental subject Final MEP pathway step enzyme; localized to plastids and proposed to be under bidirectional promoter regulation with MpHDS.
Mp5g09870 MpHDS experimental subject Late MEP pathway enzyme; localized to plastids and suggested to share bidirectional promoter regulation with MpHDR.
MpHMGR missing experimental subject Rate-limiting MVA pathway enzyme; localized to the ER and Golgi in oil body cells. A truncated feedback-insensitive form (Mp-tHMGR) was also tested for boosting precursor flux.
Mp3g01340 MpMVD experimental subject Mevalonate diphosphate decarboxylase reported with oil body-specific expression/localization in this study.
Mp4g01910 MpSQS3 experimental subject Squalene synthase reported as broadly expressed and associated with sterol biosynthesis; its localization was examined in this study.
Mp5g23030 MpACT2 sequence background Acetoacetyl-CoA thiolase (MVA pathway) listed in the gene table with predicted peroxisomal/soluble localization; not experimentally characterized.
Mp3g02320 MpC1HDZ citation background Cited transcription factor implicated in oil body cell differentiation from prior work.
Mp6g08690 MpERF13 citation background Cited as a transcription factor involved in oil body cell differentiation from prior developmental work.
Mp4g22640 MpHMGS sequence background MVA pathway enzyme listed among annotated terpene biosynthesis genes; not experimentally characterized here.
Mp4g00610 MpIDI1 sequence background Isopentenyl diphosphate isomerase listed among annotated terpene pathway genes; not experimentally characterized.
Mp6g12890 MpIDI2 sequence background Second isopentenyl diphosphate isomerase isoform listed in the gene tables; not experimentally characterized.
Mp3g22530 MpIDS1 sequence background Annotation corresponding to MpFPS, the farnesyl pyrophosphate synthase characterized in this study.
Mp2g13280 MpIDS2 sequence background Former annotation re-annotated in this paper as MpGGPPS (geranylgeranyl pyrophosphate synthase).
Mp5g21200 MpIDS3 sequence background Alternative isoprenyl diphosphate synthase analyzed in silico; lower expression or distinct predicted function compared with the characterized IDS genes.
Mp5g20580 MpIDS4 sequence background Alternative isoprenyl diphosphate synthase listed/analyzed with lower expression or different predicted function.
Mp1g06110 MpIDS5 sequence background Alternative isoprenyl diphosphate synthase listed/analyzed with lower expression or different predicted function.
Mp4g13410 MpMK sequence background Mevalonate kinase listed among MVA pathway enzymes in the gene tables; not experimentally characterized.
MpMYB2 missing citation background Cited transcription factor implicated in oil body cell differentiation.
Mp8g14400 MpPMK sequence background Phosphomevalonate kinase listed among MVA pathway enzymes; not experimentally characterized.
Mp2g21100 MpSQS1 sequence background Squalene synthase isoform listed among annotated genes; not experimentally characterized here.
Mp3g14160 MpSQS2 sequence background Squalene synthase isoform listed among annotated genes; not experimentally characterized here.
Mp2g13330 MpTGA citation background Cited transcription factor implicated in oil body cell differentiation.