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The phosphorylated pathway of serine biosynthesis affects sperm, embryo, and sporophyte development, and metabolism in Marchantia polymorpha.

Wang, M., et al. · 2024a · Communications Biology   research

doi:10.1038/s42003-023-05746-6   PMID:38267515   PMC10808223

Linked genes (14)  2 core 12 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp8g16970 MpPGDH missing experimental subject Main study gene; 3-phosphoglycerate dehydrogenase of the phosphorylated serine biosynthesis pathway. CRISPR/Cas9 knockouts show serine supply in the dark is crucial for thallus growth and that MpPGDH-mediated serine synthesis is essential for sperm, embryo, and sporophyte development. Recombinant protein (65.6 kDa) shows AtPGDH1-like cooperative inhibition by L-serine.
Mp1g15430 MpPSAT missing experimental subject 3-phosphoserine aminotransferase, second enzyme of the phosphorylated pathway. Its expression together with MpPGDH was significantly induced 8 and 16 h after transfer to darkness, supporting activation of the pathway under dark conditions.
Mp6g10990 MpACT1 experimental tool Reference gene used to normalize relative expression levels in qRT-PCR.
Mp1g13010 MpDUO1 experimental tool Sperm differentiation marker (upstream regulator of MpPRM); its expression was similar in wild type and Mppgdh mutants in the qRT-PCR panel, distinguishing affected from unaffected steps.
Mp3g14390 MpPRM experimental tool Late-spermiogenesis sperm-specific marker; significantly repressed in Mppgdh mutants in the qRT-PCR sperm-differentiation panel, used as a readout of impaired sperm development.
Mp1g00710 MpCEN1 sequence background Listed only in the data availability/accession section (Mp1g00710); no functional analysis.
Mp4g11380 MpDAZ1 sequence background Listed only in the data availability/accession section (Mp4g11380); no functional analysis.
Mp2g07580 MpGDH missing experimental background Glycerate dehydrogenase of the glycerate pathway; qRT-PCR shows gradual downregulation in darkness as an alternative serine route.
Mp6g01560 MpLC7 sequence background Listed only in the data availability/accession section (Mp6g01560); no functional analysis.
Mp1g11830 MpMID sequence background Listed only in the data availability/accession section (Mp1g11830); no functional analysis.
Mp2g10500 MpPSP missing sequence background 3-phosphoserine phosphatase, named as the third enzyme of the phosphorylated pathway; no experimental functional data presented.
Mp3g04030 MpRKD sequence background Listed only in the data availability/accession section (Mp3g04030); no functional analysis.
Mp1g09830 MpSHMT missing experimental background Serine hydroxymethyltransferase of the glycolate pathway; measured by qRT-PCR and gradually reduced in darkness, cited as an alternative serine route.
Mp4g08430 MpTUA5 sequence background Listed only in the data availability/accession section (Mp4g08430); no functional analysis.