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Efficient CRISPR/Cas9-based genome editing and its application to conditional genetic analysis in Marchantia polymorpha.

Sugano, S. S., et al. · 2018 · PLoS ONE   research

doi:10.1371/journal.pone.0205117   PMID:30379827   PMC6209168

Linked genes (7)  3 core 4 peripheral

Gene ID Name Evidence / Role Function (this paper)
Mp1g12750 MpARF1 experimental subject Primary target for evaluating CRISPR/Cas9 editing efficiency; Mparf1 mutants show NAA-resistant phenotype enabling positive selection. Co-transformation yielded hundreds of NAA-resistant T1 plants, all 16 sequenced harbored indels/substitutions, with no off-target mutations at three predicted sites.
Mp2g08950 MpMPK1 experimental subject Essential MAPK gene (knockouts unobtainable by gene targeting) studied via conditional knockout; CRISPR alone gave only mosaic mutants while co-transformation with cDNA complementation vectors yielded monoclonal frameshift mutants. Heat-shock/dexamethasone-induced Cre/loxP excision of the rescuing cDNA caused severe growth defects, confirming a critical role in growth and development.
Mp8g11800 MpNOP1 experimental subject Targeted to assess editing efficiency via the easily scored transparent-thallus (air-chamber-less) phenotype; 87.5% of 32 T1 plants were mutated. Used to test guide length (17nt or fewer dramatically less efficient) and to induce 0.5/1.5/4.5kb deletions with paired gRNAs; no off-target mutations.
Mp3g23400 MpEF1 experimental tool MpEF promoter drives Atco-Cas9 expression in editing vectors; preferentially expressed in meristematic tissues, immunoblot showed higher Cas9 accumulation than hCas9.
MpHSP17.8A1 missing experimental tool MpHSP17.8A1 heat-shock promoter drives Cre-GR for heat-shock/dexamethasone-inducible conditional recombination in the knockout system.
Mp5g03810 MpPHOT experimental tool Mpphot (phototropin) detected by antibody as loading control in immunoblot analysis of Cas9 expression.
MpU6-1 missing experimental tool MpU6-1 promoter (2kbp and 500bp versions) drives gRNA expression; both lengths gave comparable genome-editing efficiency.