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Normal oil body formation in Marchantia polymorpha requires functional coat protein complex I proteins.

Kanazawa, T., et al. · 2022 · Frontiers in Plant Science   research

doi:10.3389/fpls.2022.979066   PMID:36046592   PMC9420845

Linked genes (13)  3 core 10 peripheral

Gene ID Name Evidence / Role Function (this paper)
MpRET1 missing experimental subject Alpha COPI subunit; tested by beta-estradiol-inducible knockdown with significantly reduced mRNA. Knockdown caused abnormally shaped oil bodies and severe developmental inhibition, consistent with COPI dysfunction in oil body formation.
MpSEC21 missing experimental subject Gamma COPI subunit; tested by beta-estradiol-inducible knockdown with qRT-PCR-confirmed reduced mRNA. Knockdown produced abnormally shaped oil bodies and severely impaired growth (more severe than loss of MpSEC28), supporting a COPI role in oil body morphogenesis.
Mp1g28220 MpSEC28 missing experimental subject Causal gene of the screened T-DNA mutant; encodes the epsilon COPI subunit (T-DNA in 5' UTR of Mp1g28220). Mpsec28 mutants show rounded oil bodies (circularity 0.82 vs WT 0.45). MpSEC28-mCherry forms ring-shaped structures at the Golgi periphery, colocalizing with COPI markers.
Mp3g25140 MpAPT experimental tool Used as the internal reference gene for qRT-PCR normalization of target transcript levels.
MpERD2 missing experimental tool Alternative Golgi marker (MpERD2-mCitrine); ring-like MpSEC28 structures associated with the Golgi periphery labeled by it.
Mp1g19070 MpGOS11 experimental tool Golgi apparatus marker (mCitrine-MpGOS11); used for colocalization showing MpSEC28-mCherry localizes to COPI-coated vesicles at the Golgi periphery.
MpIDH1 missing experimental tool Mitochondrial marker (MpIDH1-mCitrine) used as negative control; low colocalization with MpSEC28 confirmed absence from mitochondria.
Mp1g08940 MpRAB5 experimental tool Multivesicular body marker (mCitrine-MpRAB5); no clear colocalization with MpSEC28, excluding MVB localization.
Mp4g20670 MpSYP12B experimental tool Qa-SNARE targeted to the oil-body membrane; mGFP-MpSYP12B lines used as the oil-body membrane marker to observe oil body morphology in mutants and knockdowns.
Mp2g14140 MpSYP4 experimental tool Trans-Golgi network marker (mCitrine-MpSYP4); low Pearson colocalization with MpSEC28-mCherry indicated MpSEC28 is not at the TGN.
Mp3g02320 MpC1HDZ citation background Transcription factor that, together with MpERF13, mediates oil body cell differentiation and oil body development in M. polymorpha; upstream regulator of the process studied.
Mp6g08690 MpERF13 citation background Transcription factor mediating oil body cell differentiation and oil body development in M. polymorpha; cited as upstream regulatory machinery.
Mp2g19600 MpSYP13B citation background Plasma-membrane-resident Qa-SNARE homologous to MpSYP12B that mediates secretion to the extracellular space, contrasting with MpSYP12B redirection to the oil body.